Faculty Profiles

Frandsen, E., Schwartz, B. F., Hutchins, B., Nowlin, W. H., Sams, M. D., Determan, K., … Herman, B. (2026). Relationships between benthic and hyporheic insect communities across environmental gradients in Texas Rivers.

Coleman, W., Bell, K. L., Gibson, J. R., Schwartz, B. F., Nowlin, W. H., & Nice, C. (2025). Evolutionary isolation and ecological stasis in the endangered Comal Springs Dryopid beetle (Stygoparnus comalensis). Aquatic Conservation: Marine and Freshwater Ecosystems.

Riggins, C., Perkin, J., Nowlin, W. H., & Smith, K. (2025). Population ecology and life history of Macrobrachium carcinus (Linnaeus, 1758) (Decapoda: Caridea: Palaemonidae), in a spring‐fed, headwaters river of Central Texas, USA. Journal of Crustacean Biology, 45, 1–10.

Wright, K. L., Nair, P., Swannack, T. M., & Nowlin, W. H. (2025). Assessing drivers of spatial variation in macroinvertebrate community structure in spring-fed subtropical rivers. Aquatic Ecology, https://doi.org/10.1007/s10452-025-10224-8, 1–22.

Frandsen, E., Schwartz, B. F., Hutchins, B., Nowlin, W. H., Swannack, T. M., Sams, M. D., … Cline, C. (2025). Relationships between benthic and hyporheic insect communities across environmental gradients in Texas rivers.

Thalhuber, T., Chumchal, M., Drenner, R., Nowlin, W. H., Williams, D., Barst, B., … Zudock, W. (2025). Diet and methyl mercury contamination of nestling red-winged blackbirds. Environmental Toxicology and Chemistry, 44(1), 59–67. https://doi.org/https://doi.org/10.1093/etojnl/vgae018

Perez-Rocha, M., Cottenie, K., Cline, K., Elkins, L., Mangold, R., Nowlin, W. H., … Schwalb, A. N. (2024). Using a nested sampling design across spatial scales to gain insights into distribution patterns of fishes, mussels, and macroinvertebrates in a riverine system. Journal of Biogeography, https://doi.org/10.1111/jbi.15003.

Santee, N., Conway, K., Nowlin, W. H., Smith, D., & Perkin, J. (2024). Alterations to water quality and quantity elicit similar stream fish functional trait responses in three North American rivers. Ecological Indicators, 169, 112917.

Nowlin, W. H., Stehle, M., Swannack, T. M., & Schwartz, B. F. (2024). To what extent do temperature and light availability affect metabolism of a groundwater-dominated river?

Perez Rocha, M., Austin-Bingamon, E., Sams, M. D., Santee, N., Schwartz, B. F., Perkin, J., … Schwalb, A. N. (2024). Hydrological Gradients Affect Facets of Biodiversity in Different Ways Across Distinct Organism Groups.

Perkin, J., Elkins, L., Mangold, R., Wolff, J., Perez Rocha, M., Schwalb, A. N., … Smith, D. (2024). Integrating Climate and Land Use Projections to Assess Ecological Futures for Stream Fish Assemblages Arranged Along an Aridity Gradient.

Sams, M. D., Perez Rocha, M., Schwartz, B. F., Johansen, R., & Nowlin, W. H. (2024). Patterns of Macroinvertebrate Diversity and Composition in a Spatially Complex River Basin: Taxonomic and Functional Approaches.

Determan, K., Yarnall, A., Perez Rocha, M., Schwartz, B. F., & Nowlin, W. H. (2024). Macroinvertebrate Community Responses Along Climate and Disturbance Gradients: Testing Predictions of the Stream Biome Gradient Concept.

Nair, P. R., Gibson, J. R., Schwartz, B. F., & Nowlin, W. H. (2023). Temperature responses vary between riffle beetles from contrasting aquatic environments. Journal of Thermal Biology, 112, 103485.

Worsham, W., Nair, P. R., Gibson, R., Rodriguez, D., Schwartz, B. F., & Nowlin, W. H. (2023). Phylogenetically independent behavior mediating geographic distributions suggests habitat is a strong driver of phenotype in crangonyctid amphipods. Frontiers in Ecology and Evolution, 11, doi:10.3389/fevo.2023.1234244.

Bowles, D., & Nowlin, W. H. (2023). Ecology of Aquatic Invertebrates in Springs and Headwater Streams: Imperiled Habitats in a Changing World. Hydrobiology, 2(3), 463–466.

Saltus, C., Johansen, R., Reif, M., Nowlin, W. H., Schwartz, B. F., & Perkin, J. (2023). Next Generation Ecological Models-Central Texas Watersheds: Geospatial Layers and Related Tables. Engineer Research and Development Center (US). https://doi.org/http://dx.doi.org/10.21079/11681/47608

Perez, K. E., Noreika, N., Norris, C., Kelly, M., Lopez, M., Ortega, C., … Nowlin, W. H. (2022). Population density and reproductive seasonality of Tryonia cheatumi (Gastropoda: Cochliopidae), the Phantom Tryonia. Freshwater Mollusk Biology and Conservation, 25, 82–90.

Bassett, L. G., Mali, I., Nowlin, W. H., Foley, D. H., & Forstner, Mi. R. (2022). Diet and Isotopic Niche of the Rio Grande Cooter (Pseudemys gorzugi) and Syntopic Red-eared Slider (Trachemys scripta elegans) in San Felipe Creek, Texas, USA. Chelonian Conservation and Biology, 21(2), 199–211.

Coleman, W., Nice, C., Nowlin, W. H., & Schwartz, B. F. (2022). Population Genetics and Trophic Ecology of the Comal Springs Dryopid Beetle.

Gibson, R., Barr, C., Diaz, P., Schwartz, B. F., & Nowlin, W. H. (2022). Recent discoveries of diverse groundwater stygobiontic riffle beetles in Texas.

Perkin, J., Elkins, L., Marigold, R., Perez Rocha, M., Schwalb, A. N., Schwartz, B. F., … Smith, D. (2022). Existing Aridity Gradient Mirrors Future Fish Assemblage Projections Under Climate Change Scenarios.

McCormack, M. A., Nowlin, W. H., & Dutton, J. (2021). Effect of trophic position on  mercury concentrations in bottlenose dolphins (Tursiops trancatus) from the Northern Gulf of Mexico. Environmental Research, 204, 112–124.

Nair, P., Diaz, P. H., & Nowlin, W. H. (2021). Interactions at surface – subterranean ecotones: structure and function of food webs within spring orifices. Oecologia, 196, 235–248.

Rodibaugh, K. R., Becker, J. C., Timmins, G., & Nowlin, W. H. (2020). Physicochemical and carbon quantity–quality gradients equally influence bacterial carbon metabolism across an arid riverscape. Aquatic Ecology, 54, 677–696.

Perez, K., Noreika, N., Norris, C., Kelly, M., Lopez, M., Ortega, C., … Nowlin, W. H. (n.d.). Population density and reproductive seasonality of Tryonia cheatumi (Gastropoda; Cochliopidae), the phantom tryonia. Freshwater Mollusk Biology and Conservation.

Nair, P. R., Huertas Pau, M. del M., & Nowlin, W. H. (2020). Metabolic responses to long-term food deprivation in subterranean and surface amphipods. Subterranean Biology, 33, 1–15. https://doi.org/10.3897/subtbiol.33.48483

Zawalski, R., Nowlin, W., Cottenie, K., Grubh, A., & Schwalb, A. N. (2019). Distinctive macroinvertebrate communities in a subtropical river network. Freshwater Ecology, 34, 135–150.

Ortega-Rodriguez, C., Chumchal, M., Drenner, R., Kennedy, J., Nowlin, W., Barst, B., … Basu, N. (2019). Relationship between methyl mercury contamination and the proportion of aquatic and terrestrial prey in diets of seven taxa of shoreline spiders. Environmental Toxicology and Chemistry, 38, 2503–2508.

Nair, P., Worsham, M., Hunter, A., Gibson, R., & Nowlin, W. H. (2019). Sexual dimorphism in three species of Heterelmis Sharp (Coleoptera:  Elimidae), 73, 1075–1083.

Jeong, J., Wolfe, III, J., Schwartz, B. F., Loiacomo, D., Nowlin, W., & Furl, C. (2019). Time-scale effects upon SWAT simulations investigating urban watershed sediment export. Retrieved from https://swat.tamu.edu/conferences/2019-vienna/agenda/

Schwartz, B. F., Nowlin, W. H., Hardy, T., Jeong, J., & Wolfe, J. (2019). Sessom Creek Sediment Export Study. Texas State University.

Olson, J., Robertson, J., Swannack, T. M., McMahon, R. F., Nowlin, W. H., & Schwalb, A. N. (2018). Dispersal of zebra mussels (Dreissena polymorpha) downstream of an invaded lake. Aquatic Invasions, 13, 199–209.

Loiacomo, D., Schwartz, B. F., & Nowlin, W. H. (2018). Stormwater and Non-Point Source Contaminants in a Small Urban Watershed. https://doi.org/doi: 10.1130/abs/2018AM-323561

Loiacomo, D., Schwartz, B. F., Nowlin, W. H., & Timmins, G. (2018). Stormwater and Non-Point Source Contaminants in Sessom Creek, San Marcos.

Loiacomo, D., Schwartz, B. F., Nowlin, W. H., & Timmins, G. (2018). Stormwater And Non-Point Source Contaminants in Sessom Creek, San Marcos, TX.

Becker, J. C., Rodibaugh, K. J., Hahn, D., & Nowlin, W. H. (2017). Bacterial community composition and carbon metabolism in a subtropical riverscape. Hydrobiologia, 792(1), 209–226.

Zawalski, R., Nowlin, W., Hardy, T., Cottenie, K., & Schwalb, A. N. (n.d.). Developing habitat suitability criteria for benthic macroinvertebrates in the lower Guadalupe River - Final Report. Texas Parks and Wildlife Department.

Coleman, W. T., Nice, C., Nowlin, W. H., & Schwartz, B. F. (2017). Cryptic, sympatric diversity uncovered in Lirceolus isopods at a stygobiont hot-spot, Edwards Aquifer, Texas.

Nowlin, W. H., Hutchins, B., Engel, A., & Schwartz, B. (2016). Chemolithoautotrophy supports subterranean food webs and affects diversity and stability in groundwater communities. Ecology, 97, 1530–1542.

Hardy, T., Crawford, M., Agare, O., Williamson, P. S., & Nowlin, W. H. (2016). Suspended sediment impacts on Texas wild-rice and other aquatic plant growth characteristics and aquatic macroinvertebrates.

Nowlin, W. H., Diaz, P., Fries, J., & Alexander, M. (2015). Mesohabitat associations of the threatened San Marcos salamander (Eurycea nana) across its geographic range. Aquatic Conservation: Marine and Freshwater Ecosystems.

Nowlin, W. H., Datri, C., Pray, C., & Zhang, Y. (2015). Nutrient enrichment scarcely affects ecosystem impacts of a non-native herbivore in a spring-fed river. Freshwater Biology.

Nowlin, W. H., Diaz, P., & Forstner, M. (2015). Survey of aquatic macroinvertebrates in a National Wetland are in Bastrop County. Texas Journal of Science.

Nowlin, W. H., Hutchins, B., & Schwartz, B. (2014). Morphological and trophic specialization in a subterranean amphipod assemblage. Freshwater Biology.

Nowlin, W. H., Becker, J., Rodibaugh, K., Labay, B., Bonner, T., & Zhang, Y. (2014). Physiographic gradients determine nutrient concentrations more than land use in a Gulf Slope (USA) system. Freshwater Science.

Nowlin, W. H., Brown, D., Ozel, E., Mali, I., Episcopo, D., Jones, M., & Forstner, M. (2013). Comparison of short term low, moderate, and high severity fire impacts to aquatic and terrestrial ecosystem components of a southern USA mixed pine/hardwood forest. Forest Ecology and Management.

Nowlin, W. H., Capps, K., Atkinson, C., Rugetski, A., Boersma, K., Carey, C., … Vaughn, C. (2012). Impacts of species addition and species loss on ecosystem function in freshwater systems. Bulletin of the Ecological Society of America.

Jones, T., Chumchal, M., Drenner, R., Nowlin, W. H., & Timmins, G. (2012). Bottom-up nutrient and top-down fish impacts on contaminant flux from aquatic ecosystems. Environmental Toxicology and Chemistry.

Nowlin, W. H., Brown, D., DeVolld, B., & Forstner, M. (2012). Ants in Houston toad habitat: annual activity and responses to canopy cover and fire. Journal of Fish and Wildlife Management.

Nowlin, W. H., Scott, S., Pray, C., & Zhang, Y. (2012). Effects of native and introduced consumers on stream ecosystem function. Aquatic Sciences.

Nowlin, W. H., Henderson, B., Chumchal, M., Drenner, R., & Diaz, P. (2012). Mercury contamination of macroinvertebrates from grassland ponds with and without fish. Environmental Toxicology and Chemistry.

Becker, J., Nowlin, W. H., Groeger, A., Chumchal, M., & Hahn, D. (2011). Spatial variability in the speciation and bioaccumulation of mercury in an arid subtropical reservoir ecosystem. Environmental Toxicology and Chemistry.

Nowlin, W. H., Pound, K., Bonner, T., & DG Huffman, D. (2011). Trophic ecology of a nonnative population of suckermouth catfishes (Hypostomus) in a central Texas spring-fed stream. Environmental Biology of Fishes.

Nowlin, W. H., Hardy, T., Schwartz, B., & Bonner, T. (2011). Implementation of the San Marcos River Observing System: a Spatially Explicit Delineation of a Spring System based on Physical, Hydrologic, Chemical, and Biological Monitoring.

Nowlin, W. H., Davies, J., Matthews, B., & Mazumder, A. (2010). Temporal discontinuity of nutrient limitation in plankton communities. Aquatic Sciences.

Nowlin, W. H., Smith, A., Abuzeineh, A., & Bonner, T. (2010). Mercury contamination of the fish community of a semi-arid and arid river system: Spatial variation and the influence of environmental gradients. Environmental Toxicology and Chemistry.

Nowlin, W. H., Caston, C., Gaulke, A., & Vanni, M. (2009). The relative importance of heterotrophic bacteria to pelagic ecosystem dynamics varies with reservoir trophic state. Limnology and Oceanography.

Nowlin, W. H., Pray, C., & Vanni, M. (2009). Deposition and decomposition of periodical cicadas (Homoptera: Cicadidae) in woodland aquatic systems. Journal of the North American Benthological Society.

Nowlin, W. H., Vanni, M., & Yang, L. (2008). Comparing resource pulses in aquatic and terrestrial systems. Ecology.

Nowlin, W. H., González, M., Stevens, M., Vanni, M., Fields, M., & Valente, J. (2007). Allochthonous subsidy of periodical cicadas affects dynamics and stability of pond communities. Ecology.

Nowlin, W. H., Davies, J.-M., & Mazumder, A. (2007). Planktonic phosphorus pool sizes and cycling efficiency in coastal and interior British Columbia lakes. Freshwater Biology.

Nowlin, W. H., Drenner, R., Guckenberger, K., Lauden, M., Alonso, T., Fennell, J., & Smith, J. (2006). Gape limitation, prey size refuges and the top-down impacts of piscivorous largemouth bass in shallow pond ecosystems. Hydrobiologia.

Nowlin, W. H., Chumchal, M., Drenner, R., & Mann, S. (2005). Biomass-dependent effects of common carp on water quality in shallow ponds. Hydrobiologia.

Nowlin, W. H., Evarts, J., & Vanni, M. (2005). Release rates and potential fates of nitrogen and phosphorus from sediments in a eutrophic reservoir. Freshwater Biology.

Nowlin, W. H., Davies, J.-M., & Mazumder, A. (2004). Temporal changes in nitrogen and phosphorus co-deficiency in plankton in lakes of coastal and interior British Columbia. Canadian Journal of Fisheries and Aquatic Sciences.

Nowlin, W. H., Davies, J.-M., Nordin, R., & Mazumder, A. (2004). Effects of water level fluctuation and short-term climate variation on thermal and stratification regimes of a British Columbia reservoir and lake. Lake and Reservoir Management.

Nowlin, W. H., Davies, J.-M., & Mazumder, A. (2004). Variation in temporal [14C] plankton photosynthesis among warm monomictic lakes of coastal British Columbia. Journal of Plankton Research.

Nowlin, W. H., Spafard, M., Davies, J.-M., & Mazumder, A. (2002). A morphometric atlas of selected lakes in southern British Columbia: Vancouver Island, Saltspring Island and the Kooteney region.

Nowlin, W. H., & Drenner, R. (2000). Context-dependent effects of bluegill on experimental mesocosm communities. Oecologia.

Nowlin, W. H., Becker, J., Rodibaugh, K., & Hahn, D. (1925). Bacterial carbon metabolism and biogeography in a sub-tropical riverscape. Hydrobiologia.